. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. 136 Steven M. Goodman et al.: A new species of Eliunts from the Réserve Speciale d'Ankarana, northern Madagascar rithm. For ML analyses, we first identified the best-fit model of nucleotide substitution using the Akaike infor- mation criterion (AIC) as employed in ModelTest ver. (Posada & Crandall 1998). We subsequently evaluat- ed whether a molecular clock fit our data using a hierar- chical log-likelihood ratio test. Parameters for the result- ing best-fit
. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. 136 Steven M. Goodman et al.: A new species of Eliunts from the Réserve Speciale d'Ankarana, northern Madagascar rithm. For ML analyses, we first identified the best-fit model of nucleotide substitution using the Akaike infor- mation criterion (AIC) as employed in ModelTest ver. (Posada & Crandall 1998). We subsequently evaluat- ed whether a molecular clock fit our data using a hierar- chical log-likelihood ratio test. Parameters for the result- ing best-fit model were fixed in a heuristic search using 10 replicates of random taxon addition and TBR branch swapping. Nodal support was calculated for both MP and ML analyses using non-parametric bootstrapping (Felsen- STEIN 1985). All bootstrap analyses employed 1000 pseudoreplicates analyzed with heuristic searches as above. Polymorphism and divergence statistics were cal- culated using DNAsp ver. (Rozas et al. 2003); in ad- dition, we report ML-corrected divergence values as cal- culated in PAUP*. 3. RESULTS After comparisons of the specimens from the Réserve Spé- ciale (RS) d'Ankarana, previously identified as EUurus antsingy, using external and craniodental measurements and characters, as well as a molecular analysis, these an- imals represent a previously unknown species of Mala- gasy rodent falling within the E. antsingy group (sensu Carleton & Goodman 2007). This new taxon is de- scribed below. Superfamily Muroidea Illiger, 1811 Family Nesomyidae Major, 1897 Subfamily Nesomyinae Major, 1897 EUurus carletoni, new species Figs. 2-3; Tables 1-3 Holotype. An adult female specimen in the Field Muse- um of Natural Histoi7(FMNH 173105) prepared as skin, skull, and partial skeleton, collected 9 April 2002 by Steven M. Goodman (original number SMG 12832). The round skin and skull are in fine condition. Recorded ex- ternal data include: TOTL, 335 mm; HBL, 143 mm; TL, 183 mm;
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