. Biology of the Heteromyidae. Heteromyidae. TROPICAL ECOLOGY 605 PANAMA - R. 180 3 3 Fig. 2.—Continued. peared not to hold at Panama-S where im- matures were relatively numerous late in the rainy season when low levels of fruitfall were observed (Fleming, 1971: table 18). Survivorship differed among populations of H. desmarestianus. At La Selva, individuals lived longer and population annual survival was higher than at Los Tuxtlas (Table 2). In both locations, however, most individuals that were trapped regularly disappeared from the grid after one year (Fleming, 1974a; Sanchez-Cordero, in pr
. Biology of the Heteromyidae. Heteromyidae. TROPICAL ECOLOGY 605 PANAMA - R. 180 3 3 Fig. 2.—Continued. peared not to hold at Panama-S where im- matures were relatively numerous late in the rainy season when low levels of fruitfall were observed (Fleming, 1971: table 18). Survivorship differed among populations of H. desmarestianus. At La Selva, individuals lived longer and population annual survival was higher than at Los Tuxtlas (Table 2). In both locations, however, most individuals that were trapped regularly disappeared from the grid after one year (Fleming, 1974a; Sanchez-Cordero, in press a). No data on survivorship of adults or immatures are available at Monteverde or Panama-S for comparison. Liomys densities fluctuated seasonally and appeared to be related to the rainfall pat- tern. Population density of Liomys salvini at La Pacifica ranged from four individuals/ ha in the dry season to eight individuals/ha in the rainy season in 1971 (Fig. 1). A high proportion of adults was present in the dry season, and most immatures were captured in the rainy season (Fleming, 1974a). At Panama-R, L. adspersus density averaged 10 individuals/ha during the 1966 rainy season, declining to five individuals/ha dur- ing the dry season (Fig. 1). Recruitment of juveniles began in the dry season and peaked early in the rainy season of 1967 (Fleming, 1971). L. salvini and L. adspersus densities in- creased early in the rainy season and then decreased as the rainy season progressed and during the dry season. No information is available for evaluating year-to-year de- mographic variation in tropical dry forests, except for the limited information from L. salvini. Fleming (1974a) observed that pop- ulation levels appear to be stable between years at La Pacifica, but D. Janzen (pers. comm.) has found considerable year-to-year variation in L. salvini density at Santa Rosa National Park in Costa Rica. Certainly, more data on this topic are badly needed to de- termine the effects of
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