. The Canadian field-naturalist. 1993 Goodchild: Status of the Tessellated Darter 427. Figure 4. Canadian distribution of the Tessellated Darter, Etheostoma olmstedi. Habitat Tessellated Darters are tolerant of a wide range of habitat characteristics. Although commonly associat- ed with pools of streams and low gradient rivers, they are also found in larger rivers and lakes (Lee and McAlUster 1980; Chapleau and Pageau 1985). The Tessellated Darter occurs in both flowing and standing waters but it shows a preference for quieter areas with slow current or still water and avoids strong riffles (S
. The Canadian field-naturalist. 1993 Goodchild: Status of the Tessellated Darter 427. Figure 4. Canadian distribution of the Tessellated Darter, Etheostoma olmstedi. Habitat Tessellated Darters are tolerant of a wide range of habitat characteristics. Although commonly associat- ed with pools of streams and low gradient rivers, they are also found in larger rivers and lakes (Lee and McAlUster 1980; Chapleau and Pageau 1985). The Tessellated Darter occurs in both flowing and standing waters but it shows a preference for quieter areas with slow current or still water and avoids strong riffles (Smith 1985). The Tessellated Darter favours shallow areas with sand, mud or rubble sub- strate. It is occasionally found in brackish waters as well (Lee and McAllister 1980; Page 1983). Physical conditions of streams occupied by study populations of Tessellated Darter in Maryland are as follows: width 6 to 15 m (20-50 feet), depth 13 cm (5 inches) in riffles to nearly a meter (3 feet) in pools (Tsai 1972). McAllister et al. (1972) located the Tessellated Darter in larger rivers (over 30 m wide) in the Ottawa area, in particular the Mississippi, Rideau, and Ottawa rivers. By contrast, the Johnny Darter is usually established in smaller streams (less than 15 m), although it also is found in the Ottawa River. General Biology Reproductive Capability Studies designed to investigate the life history of the Tessellated Darter have not been conducted in Canada. The following information is gathered from published studies in the United States. The Tessellated Darter spawns in late spring. The exact time is dependent upon a combination of suit- able temperature and optimal photoperiod. Spawning has been reported from April to June throughout its range with yearly fluctuations due to varying water temperature (Raney and Lachner 1943; Lee and McAllister 1980; Page 1983). Tsai (1972) studied two isolated populations of Tessellated Darters in the Patuxent River, Maryland. He found spawning to
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