. Bulletin of the Natural Histort Museum. Geology series. BARYONYX WALKERI 55. Fig. 44 Baryonyx walkeri, holotype, BMNH R9951; reconstruction of entire skeleton, x of this inadequate basis and because of their brevity, scarcely merit discussion. Nevertheless the authors concerned (including ourselves) have generally considered Baryonyx to be a close or not-so-close relative of Spinosaurus, so that the two genera have often been placed together as a single family (Spinosauridae Stromer, 1915). This has led certain authors (notably Sereno et al. 1994) to employ that family as a single uni


. Bulletin of the Natural Histort Museum. Geology series. BARYONYX WALKERI 55. Fig. 44 Baryonyx walkeri, holotype, BMNH R9951; reconstruction of entire skeleton, x of this inadequate basis and because of their brevity, scarcely merit discussion. Nevertheless the authors concerned (including ourselves) have generally considered Baryonyx to be a close or not-so-close relative of Spinosaurus, so that the two genera have often been placed together as a single family (Spinosauridae Stromer, 1915). This has led certain authors (notably Sereno et al. 1994) to employ that family as a single unit for the purpose of character distribution analysis. Bonaparte (1991: 18) suggested a relationship between the Spinosauridae, later fragmentary theropods from Africa (Carcharodontosaurus and Bahariasaiirus), and the South Ameri- can Abelisauridae; he claimed that the femur of all these forms possessed an anteromedially directed head and a plesiomorphically low lesser trochanter and, if this were true, then they would together represent a major diverse clade (named Neoceratosauria by Novas in 1991), distributed mainly in Gondwana. Holtz (1994a: 1105) cited i Bonaparte's work, but misleadingly stated that the Spinosauridae comprised Spinosaurus and Baryonyx; the latter genus, in fact, was never mentioned by Bonaparte. Holtz also questioned Bonaparte's suggestion, quoting our claim (1990) that Baryonyx possessed the tetanuran synapomorphy of an obturator process on the ischium; we now know that to be incorrect - the stnacture is only a flange. Nevertheless, we agree with Holtz because Baryonyx has the femo- ral head directed medially, not anteromedially, and its other characters are tetanuran rather than neoceratosaurian (see below). Meanwhile Sereno et al. (1996), on the basis of new material from the Cenomanian of Morocco, have interpreted Carcharodontosaurus as a member of the Allosauroidea. Elzanowski &Wellnhofer(1992, 1993) proposed a monophyletic group of theropods cons


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