. The Biological bulletin. Biology; Zoology; Biology; Marine Biology. 42 D. JOHNSTON AND J. FREEMAN 1 = P. chabrus (n=5| 2 = L vanegatus (n=5) 3 - C maenas (n=5) 4 - P elongalus (n=5) 5 = N inlegnlons (n=5) 6 = N- tuberculosus (n=10). 4 CDA3 - 184% -8 -L CHIT Figure 6. Results of the canonical discriminant analysis are displayed lor the second (CDA 2) and third (CDA 3) canonical discriminant func- tions evaluated at group means. Group means are central to the 95% confidence ellipses. In the bottom right corner of each graph is a vector diagram for the enzymes measured. The direction and length


. The Biological bulletin. Biology; Zoology; Biology; Marine Biology. 42 D. JOHNSTON AND J. FREEMAN 1 = P. chabrus (n=5| 2 = L vanegatus (n=5) 3 - C maenas (n=5) 4 - P elongalus (n=5) 5 = N inlegnlons (n=5) 6 = N- tuberculosus (n=10). 4 CDA3 - 184% -8 -L CHIT Figure 6. Results of the canonical discriminant analysis are displayed lor the second (CDA 2) and third (CDA 3) canonical discriminant func- tions evaluated at group means. Group means are central to the 95% confidence ellipses. In the bottom right corner of each graph is a vector diagram for the enzymes measured. The direction and length for each en/yme is an indication of the association between the en/yme and the axes and can be used to interpret differences among the species. Key to vector diagram: a-GLU = a-glucosidase, AMYL = a-amylase, |3-GLU = /3-glu- cosidase. CELL = cellulase, CHIT = chitinase. LAM = laniinarmase. PROT = total protease. TRYP = trypsin. ately consistent with an omnivorous feeding strategy. Al- though it is generally accepted that high protease activity reflects a carnivorous diet, other studies have also found high proteolytic activity in omnivores (Jonas el al., 1983; Hidalgo et ul., 1999). The sea lettuce (Ulvn Icictncu). which is ingested in large quantities by L. variegatus (Lobhan and Harrison. 1997). is high in protein (15% of the organic matter) and may contribute to the high protease activity in this crab. More in line with an omnivorous diet, the a-glucosidase activity was about twice as high in L. variegatus as in any other species we studied. This high a-glucosidase activity was responsible for separating L. variegatus from all other crab species in the MANOVA and, coupled with substantial a-amylase activity, suggests that L. variegatus is well equipped to utilize the carbohydrates within its diet. The strong activity of a-enzymes indicates that a-linked storage carbohydrates are important in its diet. Such storage prod- ucts are present in both green algae (, starch, wh


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Keywords: ., bookauthorlilliefrankrat, booksubjectbiology, booksubjectzoology