. Croonian Lecture: The Respiratory Process in Muscle and the Nature of Muscular Motion . veseen already, that in oxygen the carbon dioxide yield of the muscle wasincreased threefold or more (see fig. 1) and that, nevertheless, in spite of thisincreased combustion the irritability, as many from Humboldt to Joteyko hadpreviously found, was not more quickly exhausted but longer irritant gases increase the yield of carbon dioxide by quickening theproduction of the lactic acid, which expels preformed carbon dioxide held inthe muscle. But oxygen, while it would set a combustion flari
. Croonian Lecture: The Respiratory Process in Muscle and the Nature of Muscular Motion . veseen already, that in oxygen the carbon dioxide yield of the muscle wasincreased threefold or more (see fig. 1) and that, nevertheless, in spite of thisincreased combustion the irritability, as many from Humboldt to Joteyko hadpreviously found, was not more quickly exhausted but longer irritant gases increase the yield of carbon dioxide by quickening theproduction of the lactic acid, which expels preformed carbon dioxide held inthe muscle. But oxygen, while it would set a combustion flaring, not onlydelays the stiffening of the muscle, but may altogether inhibit its onset. Amuscle forced by stimulation to stiffening may be recalled again by oxygento its previous ilaccidity (3). This seems to us to be a crucial experiment manifesting an immediate The Respiratory Process in Muscle. 451 oxidative removal of some product of activity which is a basis of fatigue andof stiffening, giving at the same time a yield of carbon dioxide as the obvioussign of a completed Hou/fs 3 f/?0M fXC/S/O/V Fig. 3.—-Changes in length of a pair of excised gastrocnemii, after fatigue. The ordinates aremeasured directly from the record upon the drum. The levers magnified 6-| times. Load3 grm. Temp. 23° C. A. Exposed to oxygen. B. Exposed to air.
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