. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. Peter Ballmann: Fossil Calidridinae from Nördlinger Ries III. Fig. 6: Left humerus of Mirolia dubia nov. spec, caudal (A), dorsal (B) and cranial (C) views. Scale 2 mm. Feature no. 3 on Fig. 8 distinguishes the mandibula from the Tringinae (Actifis, Tringa, Cataplrophoriis), while the first and the second feature are similar to Tryngites. The process is longer than in Philomachus and shorter than in Calidris and Limicola. The question whether M? mascalidris is conge


. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. Peter Ballmann: Fossil Calidridinae from Nördlinger Ries III. Fig. 6: Left humerus of Mirolia dubia nov. spec, caudal (A), dorsal (B) and cranial (C) views. Scale 2 mm. Feature no. 3 on Fig. 8 distinguishes the mandibula from the Tringinae (Actifis, Tringa, Cataplrophoriis), while the first and the second feature are similar to Tryngites. The process is longer than in Philomachus and shorter than in Calidris and Limicola. The question whether M? mascalidris is congeneric with Tiyngites siibnificoUis, can not be decided on the basis of the available fossil material. The fossil is there- fore tentatively referred to the genus Mirolia. 12. DISCUSSION OF THE RELATIONSHIPS OF MIROLIA The morphological characters of skull and bill of Miro- lia show that their closest living relatives are the short- billed, more optically oriented Calidridinae Philoma- chus and Tiyjigites. The more important features shared with these two gen- era include: - the shallow and narrow origin of m. depressor man- dibulae on the cranium and the corresponding short processus retroarticularis; - the relatively short bill, with a moderately reinforced dorsal bar. Table 8: Length measurements of Mirolia dubia nov. spec, (in mm) Skeletal element n range average s V Bill (lacrímale to apex) 26(r) Coracoid 2 Humerus 5 Ulna 4 Carpometacarpus 10 17-19 Femur 4 20 - 22 Tibiotarsus 1 37 Tarsometatarsus 6 27-30 This relationship is further supported by the postcranial features, the shape of the processus supracondylaris dorsalis on the humerus. In the number of pits in the tip of the upper jaw, Mirolia brevirostrata is close to Tiyngites. Philomachus, being of much bigger size, has a greater number of pits. Calidris comprises species of gradually differing feeding habits. The relatively short- billed forms, suc


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