. Annals of the South African Museum = Annale van die Suid-Afrikaanse Museum. Natural history. Baculites paradoxus Pervinquiere 1907 E L U '. Baculites indet. Fig. 38. Sutures of Maastrichtian Baculites paradoxus Pervinquiere, 1907, and Baculites indet. After Pervinquiere (1907, text-figs 24, 25). pattern, B. rectus, identifying unlocalized smooth Baculites is virtually impossible. In addition, some predominantly smooth species, B. yokoyamai or B. bailyi (see Klinger & Kennedy 1997) include rare forms with distinct lateral ornament. Ornamented forms present even more problems. Va


. Annals of the South African Museum = Annale van die Suid-Afrikaanse Museum. Natural history. Baculites paradoxus Pervinquiere 1907 E L U '. Baculites indet. Fig. 38. Sutures of Maastrichtian Baculites paradoxus Pervinquiere, 1907, and Baculites indet. After Pervinquiere (1907, text-figs 24, 25). pattern, B. rectus, identifying unlocalized smooth Baculites is virtually impossible. In addition, some predominantly smooth species, B. yokoyamai or B. bailyi (see Klinger & Kennedy 1997) include rare forms with distinct lateral ornament. Ornamented forms present even more problems. Variation in Eubaculites vagina, recently comprehensively figured by Klinger 1976, Kennedy & Hen- derson (1992) and Klinger & Kennedy (1993) illustrates the point—lateral ornament varies from distinctly bituberculate, through ribbed to smooth (Figs 39-41). Our material of B. capensis (Klinger & Kennedy 1997) shows a similar wide range of variation. Apart from simple intraspecific variation, similar forms of ornament appear at different times (heterochronous homoeomorphy), or in apparent unrelated baculitid lineages of more-or-less the same age (synchronous homoeomorphy). Thus the crescentic dorsolateral tubercles of B. brevicosta in the Coniacian are virtually identical to those of Maastrichtian B. lomaensis. Some Lower Campanian B. sulcatus with strong lateral and bifurcating ventral ribbing are indistinguishable from similar Upper Campanian B. leopoliensis. Specimens with identical lateral tubercles occur in B. capensis and B. codyensis, although populations differ markedly. Identical ornament consisting of distinct auricular lateral nodes occurs in the lineages of B. vanhoepeni and B. sulcatus from the Middle and Lower Campanian of Zululand and Pondoland respectively, B. asperiformis Meek, 1876, in the Middle Campanian of the Western Interior, B. taylorensis in the Middle Campanian of the Gulf Coast and, in Upper Santonian, Boehmoceras arcuius. Unlocalized specim


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