Contributions from the United States National Herbarium . pex acute,scabrous; lower glumes 2-3 mm long, 1-nerved;upper glumes 3-4 mm long, 3-nerved; lemmas 4-5 mm long, lanceolate, 5-nerved, membranousto coriaceous, purplish, upper 1/3 scabrous, apexmucronate; callus glabrous; paleas as long as thelemma, membranous, papillose, upper 1/3 hairy,lodicules lanceolate, acuminate; anthers long; ovary apex glabrous. Caryopses lanceo-late; hilum 3/5 of total lenght. Leaf blade anatomy.—Cross-sections with7-15 vascular bundles and 5-13 ribs above; scle-renchyma under abaxial epidermis discont
Contributions from the United States National Herbarium . pex acute,scabrous; lower glumes 2-3 mm long, 1-nerved;upper glumes 3-4 mm long, 3-nerved; lemmas 4-5 mm long, lanceolate, 5-nerved, membranousto coriaceous, purplish, upper 1/3 scabrous, apexmucronate; callus glabrous; paleas as long as thelemma, membranous, papillose, upper 1/3 hairy,lodicules lanceolate, acuminate; anthers long; ovary apex glabrous. Caryopses lanceo-late; hilum 3/5 of total lenght. Leaf blade anatomy.—Cross-sections with7-15 vascular bundles and 5-13 ribs above; scle-renchyma under abaxial epidermis discontinuous,sometimes abaxial continuous, extending to some(rarely to all) vascular bundles forming girders;adaxial epidermis with scattered hairs, the mm long. Observations.—Festuca toca differs from theremaining species oiF. sect. Cataphyllophorae byhaving solitary culms. Festuca toca is morphologi-cally similar to F. andicola (F sect. Aulaxyper), butdiffers by having longer glumes (lower glumes2-3 versus mm; upper glumes 3^ versus. -80 -75 -70 -65 Figure 39. Distribution of Festuca asplundii (•), F. toca (A), and F cocuyana (?). 84 Festuca in South American Paramos mm) and longer anthers ( mm). Distribution and habitat.—This species isendemic to Colombia where it is known only fromthe Cordillera Oriental (Boyaca, Norte de Santander).It occurs in swampy patches in grass paramo zonebetween 3100-3500 m. Festuca toca is present indifferent vegetation communities such as: Lysipomiasphagnophilla subsp. minor (Cleef 1981). Additional specimens examined. : Mun. Duitama, paramo Pan de Azucar,3450 m, 30 Nov 1998, D. Stancik 1418 (COL,PRC); Paramo de Rusia, 3400 m, 2 Jan 1984, Wood4481 (COL, FMB, K); 3570 m, 13 Dec 1972, Cleef7139 (COL); summit of road from Duitama toAbendanos, 3490m, 12 Feb 1972,/). Stancik2371(COL, PRC); Hda Los Tres Corrales, 5°,73°, 3490 m, Hernandez 1022A (COL);Mun. Toca, road from Pesca
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Keywords: ., bookcentury1800, bookdecade1890, booksubjectbotany, bookyear1890
