. Annals of the South African Museum = Annale van die Suid-Afrikaanse Museum. Natural history. 162 ANNALS OF THE SOUTH AFRICAN MUSEUM individuals of the genus for, as Broom notes (1935), tuskless specimens of Dicynodon grimbeeki have more lightly-built snouts than tusked specimens. Cox reports that only two of ten specimens of Kingoria at his disposal had tusks, but describes no differences between the snouts of tusked and tuskless individuals. However, in Kombuisia the snout is more delicate and the caniniform processes are proportionately smaller than in most other dicynodonts, including tus


. Annals of the South African Museum = Annale van die Suid-Afrikaanse Museum. Natural history. 162 ANNALS OF THE SOUTH AFRICAN MUSEUM individuals of the genus for, as Broom notes (1935), tuskless specimens of Dicynodon grimbeeki have more lightly-built snouts than tusked specimens. Cox reports that only two of ten specimens of Kingoria at his disposal had tusks, but describes no differences between the snouts of tusked and tuskless individuals. However, in Kombuisia the snout is more delicate and the caniniform processes are proportionately smaller than in most other dicynodonts, including tuskless specimens of Kingoria and D. grimbeeki, and it seems unlikely that any individuals of Kombuisia had tusks. The palates of Kombuisia and Kingoria are distinctive in their lack of relief. In most dicynodonts (cf. Ewer 1961; Crompton & Hotton 1967; Cluver 1970, 1971) there is a pair of strong ridges on the back of the premaxillary beak, one on either side of the midline, and the contact between the vomer and the pre- maxilla is produced into a prominent midline ridge. In Kombuisia and Kingoria the premaxillary ridges are lacking and the midline vomerine ridge is much subdued; there is, however, a low, longitudinal ridge just medial to each maxillo-premaxillary suture (Fig. 1: LR) which is not present in other dicynodonts. The punctate and rugose sculpture on the beaks of dicynodonts is generally agreed to indicate the presence, in life, of a horny covering, because of its resemblance to comparable areas in birds and chelonians. On the rostral. Fig. 1. Kombuisia frerensis. No. 430. Skull, palatal aspect: AR, articular; LR, longitudinal ridge at lateral edge of premaxilla; MR, median ridge at junction of premaxilla and vomer; PAL, palatine; PMX, maxillo-pre- maxillary suture; Q_, Please note that these images are extracted from scanned page images that may have been digitally enhanced for readability - coloration and appearance of these illustrations may not per


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