Archive image from page 89 of Cytology, with special reference to. Cytology, with special reference to the metazoan nucleus cytologywithspec00agar_0 Year: 1920 74 CYTOLOGY CHAP. merely to the fact that the chromatin is more concentrated in the former, for as it approaches the female nucleus it grows larger, and at the same time becomes looser in texture, till ultimately the two gamete nuclei are generally of the same volume and structure. Since each gamete nucleus is haploid, it is obvious that the zygote nucleus is diploid again. As we have seen, a typical spermatozoon consists of three main
Archive image from page 89 of Cytology, with special reference to. Cytology, with special reference to the metazoan nucleus cytologywithspec00agar_0 Year: 1920 74 CYTOLOGY CHAP. merely to the fact that the chromatin is more concentrated in the former, for as it approaches the female nucleus it grows larger, and at the same time becomes looser in texture, till ultimately the two gamete nuclei are generally of the same volume and structure. Since each gamete nucleus is haploid, it is obvious that the zygote nucleus is diploid again. As we have seen, a typical spermatozoon consists of three main portions—head, middle piece, and tail. The fate of the head or nucleus Fig. 33. Syngamy in Chaeiopterns pergamenlaceus. (After Mead, , 1898.) A, soon after entry of spermato- zoon ; 9 nucleus in anaphase I. ; $ aster developing ; B, 9 nucleus in anaphase II. ; C, 9 nucleus in form of karyomeres (see p. 13T) ; 9 achromatic figure has disappeared ; D, approach ; E, fusion, of gamete nuclei. and , first and second polar bodies. we have already followed. The tail apparently takes no part in fertiliza- tion. (For the fate of the chondriosomes, which are usually situated in the tail, see p. 198.) Sometimes it does not even enter the egg, being broken off from the middle piece as soon as this has penetrated into the egg (sea-urchins). When the tail does enter the egg it breaks off from the rest of spermatozoon soon after entry, and lies in the egg cytoplasm for a time, and then degenerates. There remains to be considered the middle piece, with which the centrosome, as the development of the spermatozoon showed us, is generally related. Owing to the mode of entry of the spermatozoon— head first—the middle piece is at first behind the head. After entry, however, the head and middle piece rotate so that the latter is in front of the former during its journey towards the centre of the egg. The
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