Organography of plants, especially of the archegoniatae and spermaphyta . gnum, indeed,is evidently not a primitive but agreatly altered form, as we havealready learned when consideringits germination, and as the be-haviour of its antheridia, if weaccept the statement of Leitgeb,confirms. However this may be,at any rate it is remarkable thatin the Bryophyta the formation ofstomata repeatedly appears, as forexample in Anthoceros and in thedifferent series of the Musci, andthey in every way correspond withthe stomata of the Spermophyta. The development of the apo-physis in many species of Splach
Organography of plants, especially of the archegoniatae and spermaphyta . gnum, indeed,is evidently not a primitive but agreatly altered form, as we havealready learned when consideringits germination, and as the be-haviour of its antheridia, if weaccept the statement of Leitgeb,confirms. However this may be,at any rate it is remarkable thatin the Bryophyta the formation ofstomata repeatedly appears, as forexample in Anthoceros and in thedifferent series of the Musci, andthey in every way correspond withthe stomata of the Spermophyta. The development of the apo-physis in many species of Splachnum is remarkable, especially in S. rubrumand S. luteum^ (Fig. 125, /, //), in which the apophysis grows out into anumbrella-like fringe which in its structure resembles a dorsiventral leaf, andpossesses stomata only upon the upper side. The apophysis also takes ashare indirectly in the scattering of the spores as we shall see later. OtherSplachnaceae also have the tendency to develop large apophyses—a characterwhich has resulted in the most remarkable Fig. 125. Splachnum luteutn. /, c, capsule open ; A,apophysis. I/, unopened capsule in longitudinal section ;vS, seta ; Lx, leptoxjlem ; sp, stomata on apophysis ; cl,columella; ^, peristome; As, archesporium ; /.intercellularspace, ///and IV, diagrams to illustrate the opening ofthe capsule in Splachnum ; aa, peristome, incur\ed in Til,recur\ed in IV; (^, terminal disk of columella. /, magnified3. After Hedwig. // magTiified. After Vaizey. fllajidIV, magnified. After Bryhn. * I agree with Haberlandt in reckoning the apophysis as a portion of the seta, not of the capsule. See also Vuillemin, Sur les homologies des Mousses, Bulletin de la Societe des sciences de Nancy,xix (18S6); Bunger, Beitrage zur Anatomic der Laubmooskapsel, in Botanisches Centralblatt, xlii(1S90). * See Vaizey, On the Morpholor^y of the Sporophyte of Splachnum luteunn, in Annals of Botany,V (1890-1), p. I. i6o SPOROGONIUM OF MUSCI 3. ARRANGE
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