Contributions in geographical exploration . ished as one approachedthe Volcano, but even in the upper valley large areas which wereabsolutely barren in 1915 were coming up with occasionallupine seedlings which, though so sparse and widely scatteredthat one had to search for them, were nevertheless thrivingwith every prospect that some of them would survive. Fartherdown the valley a few areas were found where similar seedlings April, U)19j Begiuiiiiios of Revegetutioji 325 had started in 1915 and, having persisted even in deej) pumicedeposits, were flowering and seeding abundantly in liHO; whil
Contributions in geographical exploration . ished as one approachedthe Volcano, but even in the upper valley large areas which wereabsolutely barren in 1915 were coming up with occasionallupine seedlings which, though so sparse and widely scatteredthat one had to search for them, were nevertheless thrivingwith every prospect that some of them would survive. Fartherdown the valley a few areas were found where similar seedlings April, U)19j Begiuiiiiios of Revegetutioji 325 had started in 1915 and, having persisted even in deej) pumicedeposits, were flowering and seeding abundantly in liHO; whilein 1917 considerable areas as far up stream as Martin Creekwere sparsely occupied by fruiting lupines, furnishing thebasis for an increasing rate of revegetation. LUPINES THE MOST EFFECTIVE PIUNEERS. While the new vegetation in the lower valley consists ofmany species of plants, in the more exposed places lupines arethe only pioneers. (See pages 324 and 335). For this rolethey are well adapted, because of their large h(^•^•^? Ilnilugraph by Robert F. Griggs A LUPINE OX THE PUMICE FLAT AT MARTIN CREEK. These plants first appeared in 1915. They were well provided with root tubercles and grew thriftily, fruiting freely in 1917. The soil is almost entirely without organic nitrogen. which lodge where smaller seeds are blown away. On germina-tion, moreover, their large supply of stored food enables themto grow into strong plants much more quickly than the otherspecies present. But their capacity of utilizing atmosphericnitrogen through their root tubercles is probably the decisivefactor, for the ash is almost devoid of nitrogenous compounds.*Lupines growing in pumice show an abundant development ofroot tubercles which must give their possessors enormousadvantages over ordinary plants in the jirocess of revegetation. 2 Shipley, J. W. The Nitrogen Content of Katmai Ash. Iaivr No. V inthis series, pages 213-223. 320 The Ohio Journal of Science [Vol. XIX, No. 6, .Jf-*ir Wk
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