. Bulletins of American paleontology. Review of the Bullia Group: Allmon 91 fossil record quality, or "palcontological complete- ness", is therefore required. For purposes of phyloge- netic analysis, palcontological completeness can be de- fined as "the proportion of the total time and space inhabited by all the individuals of all the species of the group of interest that is preserved by fossils" (Allmon, 1989). This concept encompasses three phylogenet- ically important aspects of the completeness problem (Bretsky, 1979): (1) completeness of species diversity; (2) complete
. Bulletins of American paleontology. Review of the Bullia Group: Allmon 91 fossil record quality, or "palcontological complete- ness", is therefore required. For purposes of phyloge- netic analysis, palcontological completeness can be de- fined as "the proportion of the total time and space inhabited by all the individuals of all the species of the group of interest that is preserved by fossils" (Allmon, 1989). This concept encompasses three phylogenet- ically important aspects of the completeness problem (Bretsky, 1979): (1) completeness of species diversity; (2) completeness of stratigraphic duration of known species; and (3) completeness of geographic range of both individual species and the entire group. Each of these aspects should be considered at strati- graphic and geographic scales appropriate for the taxo- nomic level and phylogenetic question of interest. For lower taxonomic levels within single basins or prov- inces, techniques of stratigraphic completeness mea- surement can be adopted to estimate palcontological completeness. In cases of higher taxa over larger strati- graphic and geographic intervals, such methods are impractical and others are required. Bretsky (1979) proposed a method for estimating completeness of species diversity for West Atlantic Tertiary lucinid bivalves. She compared the observed diversity of fossil species throughout the Tertiary to known Recent species diversity, found approximate correspondence, and concluded that this record is "dense and continuous" enough to employ the stra- tophenetic method in a phylogenetic reconstruction (Bretsky, 1979, pp. 126-127). The pattern of diversity in the fossil record of the Bullia group differs in several respects from that of the Recent species. Text-figure 21 shows the stratigraphic distribution of all species considered here to be prob- able members of the Bullia group. The number of known species existing throughout the Cenozoic is rep- resented in Text-f
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