. Bulletin of the British Museum (Natural History), Geology. LATE CAMPANIAN-MAASTRICHTIAN ECHINOIDS 147. Fig. 18 Camera lucida drawings of Goniopygus species. A, B apical disc plating of Goniopygus arabicus sp. nov.; A, BMNH EE4017; B, BMNH EE4005. C, F, apical disc plating and ambital ambulacral tuberculation of Goniopygus superbus Cotteau & Gauthier, L12680, Geology Department, Universite de Paris VI; Senonian, Derre-i-Chahr, Iran. D, E, ambital ambulacral plating, Goniopygus arabicus sp. nov.; D, BMNH EE4017; E, BMNH EE4005. G, G. arabicus sp. nov., adoral plating, ambulacrum to left, B


. Bulletin of the British Museum (Natural History), Geology. LATE CAMPANIAN-MAASTRICHTIAN ECHINOIDS 147. Fig. 18 Camera lucida drawings of Goniopygus species. A, B apical disc plating of Goniopygus arabicus sp. nov.; A, BMNH EE4017; B, BMNH EE4005. C, F, apical disc plating and ambital ambulacral tuberculation of Goniopygus superbus Cotteau & Gauthier, L12680, Geology Department, Universite de Paris VI; Senonian, Derre-i-Chahr, Iran. D, E, ambital ambulacral plating, Goniopygus arabicus sp. nov.; D, BMNH EE4017; E, BMNH EE4005. G, G. arabicus sp. nov., adoral plating, ambulacrum to left, BMNH EE3986. Scale bars = 1 mm. primary tubercle. At the ambitus interambulacral width is 42-46% of the test diameter. There are 8 plates in a column at 25 mm diameter, rising to 11 at about 40 mm test diameter (Fig. 17). Primary tubercles are stout, imperforate and non- crenulate at the ambitus, but reduce sharply in size adapically so that the top three or four tubercles are very small. At the ambitus they are surrounded by about 6 widely-spaced scro- bicular tubercles (PI. 7, fig. 6). Down the interradius there is a single column of secondary tubercles on each plate. Adorally both columns of plates reach the peristomial margin and there is no primordial plate (Fig. 18G). The peristome is slightly invaginated and is 37^14% of the test diameter in diameter (mean = 42%, SD = 2-6%, N = 7). Buccal notches are relatively small and poorly differentiated, with only a weak rim. Remarks. This species was described under the name Goniopygus superbus Cotteau & Gauthier by Kier (1972) and Ali (1989). It differs from that species in several important respects. Firstly, the apical disc of G. superbus has a larger periproctal opening and has either five, or rarely four peria- nal tubercles (PI. 6, figs 1, 2; PI. 7, fig. 2; Fig. 18C). These perianal tubercles face upwards and the adjacent portions of the genital plates are raised in the form of a very characteris- tic stellate rim. The ra


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