. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. 186 C. Denys et lineages, of a centro-internal cusp (t4) from different primitive muroid ancestors. In- deed, many of the characters of Dendromurinae are shared with Petromyscinae, Delanymyinae, Cricetomyinae, and even some Murinae of controversial position (Acomys, Uranomys, Lophuromys) (cf. Tab. 2). The parsimony analysis indicates that the sister-taxon of Leimacomys is Tatera and that it belongs to a monophyletic group including Gerbillus and Mystromys (Fig.
. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. 186 C. Denys et lineages, of a centro-internal cusp (t4) from different primitive muroid ancestors. In- deed, many of the characters of Dendromurinae are shared with Petromyscinae, Delanymyinae, Cricetomyinae, and even some Murinae of controversial position (Acomys, Uranomys, Lophuromys) (cf. Tab. 2). The parsimony analysis indicates that the sister-taxon of Leimacomys is Tatera and that it belongs to a monophyletic group including Gerbillus and Mystromys (Fig. 3). The main reason for this clade is the absence in all these taxa of some cusps (absence of Cv5 on lower M/1—2, absence of tl and t3 on upper M2—3/) found in all other Dendromurinae and Murinae. Nevertheless, Leimacomys do not show any sign of longitudinal crest nor any of the external or cranial characters of the Gerbillinae (Denys 1993). In opposition with Mystromys and the Gerbillinae, Leimacomys has a t4. The morphological cladogram illustrated by Figure 3 implies the appearance of t4 at least two times, one time in the ancestor of a large cluster including Criceto- myinae, Deomys, Murinae, Acomys- Lophuromys and six genera of Dendromurinae and in the other time, in a lineage to which belong Gerbillinae, Mystromys, and Leimacomys. A remark can be made here concerning the position of Mystromys. Considered un- til present as a Cricetinae incertae sedis (Ellerman 1941; Carleton & Musser 1984; Denys 1991), it clusters here with Gerbillinae rather than with Murinae. According to DNA-DNA hybridization data of Rice (1974), Mystromys is as distant from true Murinae as from true Cricetinae. It is probable that Mystromys derives from an African Early Miocene Please note that these images are extracted from scanned page images that may have been digitally enhanced for readability - coloration and appearance of these illustrations may not perfectly resemble the
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