. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. 328 B. Hughes Bonn, zool. Fig. 4. Distribution of West African savanna snakes: the East African invaders. Fig. 5. Latitudinal ranges of the savanna snakes of Ghana and Upper Volta: to the left snakes which reach the coast (except no. 12: Aparallactus lunulatus and perhaps no. 147: Scaphiophis albomaculatus) and are primarily snakes of Guinea savanna, to the right snakes which do not reach the coast (except 49 — Echis ocellatus) and are pri- marily snakes of


. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. 328 B. Hughes Bonn, zool. Fig. 4. Distribution of West African savanna snakes: the East African invaders. Fig. 5. Latitudinal ranges of the savanna snakes of Ghana and Upper Volta: to the left snakes which reach the coast (except no. 12: Aparallactus lunulatus and perhaps no. 147: Scaphiophis albomaculatus) and are primarily snakes of Guinea savanna, to the right snakes which do not reach the coast (except 49 — Echis ocellatus) and are pri- marily snakes of Sudan and Sahel savanna. The latter include some of the invaders and endemics whose distribution is mapped in Figs. 2 to 4. The numbers are those used in Appendix B. Key to species: 1 Afronatrix anoscopus, 6 Amblyodipsas unicolor, 12 Aparallactus lunulatus, 16 Atractaspis aterrima, 18 Atractaspis dahomeyensis, 21 Atractaspis microlepidota, 22 Atractaspis micropholis, 23 Bitis arietans, 27 Causus ma- culatus, 30 Coluber dorri, 34 Crotaphopeltis hotamboeia, 35 Dasypeltis fasciata, 38 Dasypeltis scabra, 40 Dendroaspispolylepis, 42 Dispholidus typus, 43 Dromophis line- atus, 44 Dromophis praeornatus, 48 Echis leucogaster, 49 Echis ocellatus, 53 Elapsoi- dea semiannulata, 56 Eryx muelleri, 57 Gonionotophis granti, 59 Grayia smythii, 65 Lamprophis fuliginosus, 68 Lamprophis lineatus, 69 Leptotyphlops bicolor, 11 Lepto- typhlops macrorhynchus, 78 Leptotyphlops narirostris, 81 Leptotyphlops sundevalli, 90 Lycophidion semicinctum, 92 Mehelya crossi, 95 Meizodon coronata, 96 Meizodon regularis, 100 Naja haje, 101 Naja katiensis, 102 Naja melanoleuca, 104 Naja nigricol- lis, 106 Natriciteres olivácea, 110 Philothamnus heterolepidotus, 112 Philothamnus ir- regularis, 115 Philothamnus semivariegatus, 116 Polemon neumedi, 120 Prosymna me- leagris greigerti, 121 Prosy mna m. meleagris, 121 Psammophis elegans, 128 Psamm- ophis phillipsi, 132 Psammophis sibilans, 136 Python regius, 137 Pyth


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