. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. Superspecies in wheatears 179 a I boniger leucopyga leucura monticola phillipsi oenanthe pleschanka hispa nica monacha picata lugens finschü moesta xanthoprymna deserti pileata bottae isabellina Fig. 9: A suggested phylogeny for the genus Oenanthe. Areas of particular uncertainty in- dicated by dotted lines; plausible alternatives by arrows (see text). Alternatively, the phillipsi superspecies could conceivably be closer than indicated to the pleschanka superspecies
. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. Superspecies in wheatears 179 a I boniger leucopyga leucura monticola phillipsi oenanthe pleschanka hispa nica monacha picata lugens finschü moesta xanthoprymna deserti pileata bottae isabellina Fig. 9: A suggested phylogeny for the genus Oenanthe. Areas of particular uncertainty in- dicated by dotted lines; plausible alternatives by arrows (see text). Alternatively, the phillipsi superspecies could conceivably be closer than indicated to the pleschanka superspecies. O. oenanthe resembles the pleschanka group in general plumage pattern, though not in general ground colour. They also show similar sexual dimorphisms and non-breeding and immature plumages. However, these similarities probably represent convergence, and phillipsi differs more strongly than oenanthe from the pleschanka group. O. deserti also resembles oenanthe and the pleschanka group in plumage pattern, while O. oenanthe in its browner female, im- mature and non-breeding plumages is extremely similar to females of the picata- isabellina sequence. It is possible that the phillipsi superspecies and/or pleschanka species-group are more closely linked to deserti and the picata-isabellina sequence than indicated in Fig. 9. As might be expected, these areas of greatest uncertainty occur with respect to the earliest divisions within the genus. At the opposite extreme, the most recent splits are those about which one can be most certain. O. pleschanka and hispánica have obviously diverged very recently, or at any rate did not drift far apart genetically when in isolation from one another. They might now almost be regarded as a single species. Similarly picata, lugens and finschii have been regarded as conspecific and are obviously extremely close. Indeed, the difference between subspecies of some species (e. g. O. oenanthe oenanthe and O. o. seebohmi; O. b. bottae/frenata and O. b
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