. Atoll research bulletin. Coral reefs and islands; Marine biology; Marine sciences. 60 n. 0 09/08/96 Days after scuttling 11/09/97 ââ¢â Caledonie Toho II - â¢â¢ Caissons â*â Ricaudy Figure 2. Temporal variations of fish species richness on Caledonie Toho 2, Caissons and Ricaudy reef. A new increase phase was observed during the final sampling. These species will be referred to as "secondary" species. On the Caissons, species richness increased overall during the first 193 days from 16 to 32 species (Fig. 2). After this colonization phase, species richness remained relatively stable
. Atoll research bulletin. Coral reefs and islands; Marine biology; Marine sciences. 60 n. 0 09/08/96 Days after scuttling 11/09/97 ââ¢â Caledonie Toho II - â¢â¢ Caissons â*â Ricaudy Figure 2. Temporal variations of fish species richness on Caledonie Toho 2, Caissons and Ricaudy reef. A new increase phase was observed during the final sampling. These species will be referred to as "secondary" species. On the Caissons, species richness increased overall during the first 193 days from 16 to 32 species (Fig. 2). After this colonization phase, species richness remained relatively stable at approximately between 28 and 34 species during the rest of the survey. Variations of species richness on the natural reef (Ricaudy) were relatively low (Fig. 2). On average, 45 species were recorded during the survey. Species richness was lower in winter (40 species on average), from September to October 1996 (11 to 67 days after scuttling) and from July to mid- September 1997 (343 to 398 days after scuttling). Species richness was higher during the rest of the year (50 species on average). During the survey, species richness was significantly higher on Ricaudy than on both artificial reefs (%2 proportionality test, P ). Fish density increased rapidly during the first 109 days after scuttling (Fig. 3). This increase was mainly due to the settlement of planktivorous schooling species (Clupeidae, Apogonidae and Pomacentridae) and the migration of carnivorous species (Carangidae, Lutjanidae). After this increase, density remained relatively stable (around fish/m ) with the exceptional peaks occurring between 304 and 364 days after scuttling. These high values corresponded to the recruitment of Apogonidae, Rhabdamia spp. The density of these species decreased dramatically in the last sample because recruitment had ended and because of predation. The schools of Carangidae were feeding on small fish, which ventured out of their shelters (upper works or holds). O
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